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ANADROMOUS FISHERIES
Part 5 of 6
Striped Bass Pages 71 - 83
Striped Bass Life History Notes (Continued)
It appears that reproduction was exceptionally good
in 1953 and 1954 with progressively poorer years in
1955 and 1956. Unfortunately, statistical procedures
have revealed certain discrepancies in the sampling
methods which limit the usefulness of past surveys as
indices of abundance. A new approach is being employed
which it is hoped will yield more useful data.
Experimental data indicate the fry are usually located
nearer the surface than the bottom, although a
recent series of tests designed to determine their vertical
distribution, showed the reverse to be true on at
least one occasion. Chadwick (unpublished data) conducted
tests in the summer of 1957 which showed that
the smaller fish are found in greater numbers near the
shoreline than in mid-channel. As they approach two
inches in length, they are found more evenly dispersed
throughout the channel. Evidently, even fry
less than an inch in length have some control over
their movements despite rapid tidal currents. His findings
agree with observations made by the writer in
1954 and 1955 while engaged in this work.
Juveniles. By early summer young fish are scattered
throughout all parts of the Delta and at least
as far downstream as San Pablo Bay. Apparently, all
fry are not carried into the Delta because they can be
found far up the Sacramento and San Joaquin Rivers
in the late summer and fall. It seems rather improbable
that they return upstream after having once gone
down. Juvenile fish have been seined all along the
Sacramento and San Joaquin rivers and at least as far
down as Point Wilson in San Pablo Bay.
During seining operations during the fall (October)
of 1956 and 1957, throughout the Bay and Delta, fish
with mean lengths ranging from 2.9 to 4.0 inches
were taken. The overall average is probably about 3.5
inches. There does not appear to be any perceptible
difference of growth pattern in any particular part of
the area covered. Growth ceases, or at least is greatly
diminished from October until the following March.
Juvenile fish apparently remain principally in the
Delta for two to three years before moving into San
Francisco Bay or the ocean. During this period they
tend to be gregarious, moving about in scattered
schools.
Fish up to 16 inches may be found anywhere
throughout most of the year, but certain areas seem
to be more favorable than others. Such juveniles are
almost always present in San Pablo Bay in the vicinity
of Mare Island, the Napa River, Suisun Bay, and in a
number of localities in the Delta.
Raney (1952) states: "During the first and second
years they remain in small schools or feeding groups,
but it has been observed that they exist in large schools
by the end of the second summer."
The age at which they first begin their annual migrations
between fresh and salt water has not been
positively established on the West Coast. Most fish,
it is thought, undertake them in the third year though
many unquestionably begin in the second and others
probably wait until their fourth year or later. Inherent
differences between fishes, and sexual differences, are
likely factors influencing the age at which they move
into salt water. The proportion of adults making these
annual excursions is not known; however, since large
fish are very scarce in the Delta during the summer,
it may be concluded that the great majority are involved.
Migrations. Clark (1934 and 1936) and Calhoun
(1952) are responsible for most of our present knowledge
on striped bass migrations in California. Both
have conducted tagging experiments showing the migratory
patterns. A third study, by Chadwick, was
underway at the writing of this report.
Clark reported the results of tagging 1,544 bass,
mostly small ones (mean length 11 inches), and found
that they did not move in a well defined migration
but more or less diffused out of the tagging area.
Calhoun's work was with legal sized fish (then 12
inches and over), of which he tagged more than 4,000.
He found that unlike the smaller fish, the adults did
undertake well defined seasonal migrations. Recoveries
of tagged fish by sportsmen and gill net fishermen
showed that the adults move upstream into San Pablo
Bay and Carquinez Strait in the fall, then into the
Delta in the winter, spread out and ascend the tributary
rivers in the spring, and move down to the Bay
again by early summer.
Their movement to and in the ocean is not yet
understood, although a fair number appear to enter
the ocean each year. Occasionally, good catches are
made by surf casters off San Francisco beaches. Although
stripers are seldom taken off shore, the party
boat fleet made good catches of striped bass in the
ocean during 1956. On the Atlantic Coast extensive
north-south ocean migrations are made, presumably
for feeding purposes. This phenomenon has not been
observed to occur here.
Food Habits. Scofield (1911) found that fish up to
four inches, in Napa Creek, relied on marine worms
(50 percent) crustaceans (48 percent) and small fish
(2 percent). The items are listed in Table 25.
Hatton (1940), in a collection of 76 fish ranging
from one to six inches in length taken at Martinez,
found that 69.4 percent of all stomachs contained
crustaceans. The percentage of stomachs containing
each item found is shown in Table 26.
Hatton points out that the items were found during
September and November while the water at Martinez
was brackish.
Under freshwater conditions in this area, during
the spring, the amphipods and isopods disappear, and
the small fish were found to be feeding almost exclusively
on a species of Mysidacea (Neomysis mercedis).
The writer has on several occasions while checking
the stomachs of young-of-the-year bass (2 to 4
inches) from the San Joaquin River and Suisun Bay
during the summer also found that mysid shrimp
{Neomysis mercedis) was by far the major item in the
diet. Some of the stomachs examined were simply
packed with them. Each spring and summer the River
and Delta abound with these small crustaceans.
Messrs. Fisk and McCammon of the Department,
who have studied the food habits of the white catfish
in the Delta, observed that amphipods (Corophium
spinicorne) were the most important organism utilized
by catfish. Hatton's work seems to confirm the
importance of it in the diet of small bass. Unfortunately,
data are meager on feeding habits of small
bass from the time they begin to feed until they are
through the first year.
On the East Coast, freshwater shrimp (Gammarus),
and Dipterid (chironomid) larvae were found to be
major food items.
Striped bass become piscivorous at least by thetime they reach 6 inches and perhaps earlier. Their
diet from this size on is extremely varied, and appears
to depend upon the forage available. The larger
fish appear to have a proportionately larger percentage
of fish in the diet but even the largest specimens
were found to contain crustaceans.
Bay shrimp (Crago sp.) appears to be one of the
most common items, along with Neomysis mercedis
and the small forage fishes found in the Bay, such as
smelt, herring and anchovy. Hatton (op cit) examined
224 stomachs of adult bass taken near Pittsburg
between March 13 and May 4, 1939 and found 56.6
percent to be empty (during the spawning season).
His findings are summarized in Table 27.
Johnson and Calhoun (1952) examined 387 stomachs
of adult bass collected during a period of a year.
All fish were over 12 inches in length. One group of
229 was collected between San Rafael and Martinez
during the summer and fall, while the other lot of
158 was taken from the Delta portion of the San Joaquin
River between Antioch and the mouth of Middle
River between November 1947 and June 1948.
Shrimp (Crago sp.) were the most numerous item
and comprised the largest volume of all organisms
found in the summer group. It occurred in 35 percent
of all stomachs examined and formed 53 percent
of the volume of all foods. Anchovies, the next most
important item, occurred in 11 percent of the stomachs
and comprised 39 percent of the food volume
Isopods, crabs, mysid shrimp, and other fish were
also found but none occurred more than six times
or formed more than 2 percent of the total volume.
Of this group of fish 28 percent of the stomachs examined
were empty.
Their winter sample, the one from the Delta, contained
66 individual fish (42 percent of the sample)
with empty stomachs. Neomysid shrimp occurred in
more stomachs than any other item and formed, surprisingly
enough for large fish, 20 percent of the total
volume. Small fish, however, were the most frequent
item and accounted for the greatest volume (64 peicent).
Bay shrimp (Crago sp.) accounted for 13 percent
of the volume.
Shapovalov (1936) examined the stomachs of 47
striped bass taken from the mouth of Waddell Creek,
Santa Cruz County. He found a large variety of items,
with crustaceans predominating in the small bass, and
other fish being the principal food of the larger bass.
Bass are obviously omnivorous feeders; quoting Scofield
(1931): "Practically every marine form common
to the San Francisco Bay region has been found in
their stomachs. Their food includes fishes, such as
small Pacific herring, smelt, anchovies, split-tails,
striped bass, shad, gobies, carp and perch; crustaceans
and mollusks—crabs, shrimps, periwinkles, clams; and
various other forms such as worms, copepods and
vellella."
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Citations
Calhoun, A. J.
1949 California Striped Bass Catch Records From the Party
Boat Fishery; 1938-1948. California Fish and Game,
Vol. 35, No. 4, pp. 211-253.
1950 California Angling Catch Records from Postal Card
Surveys: 1936-1948 With an Evaluation of Postal Card
Non-response. California Fish and Game, Vol. 36, No.
3, pp. 177-233.
1951 California State-Wide Angling Catch Estimates for
1949. California Fish and Game, Vol. 37, No. 1, pp.
69-75.
1952 Annual Migrations of California Striped Bass. California
Fish and Game, Vol. 38, No. 3, pp. 391-403.
1953a. State-Wide California Angling Estimates for 1951. California
Fish and Game, Vol. 39, No. 1, pp. 103-113.
1953b. Distribution of Striped Bass Fry in Relation to Major
Water Diversions. California Fish and Game, Vol. 39,
No. 3, pp. 279-299.
1957 Striped Bass Fishing Map (Revised by John E. Skinner).
California Department of Fish and Game.
Calhoun, A. J., and John E. Skinner
1954 Field Tests of Stainless Steel and Tentalum Wire with
Disk Tags on Striped Bass. California Fish and Game,
Vol. 40, No. 3, pp. 323-328.
Calhoun, A. J., and C. A. Woodhull
1948 Progress Report on Studies of Striped Bass Reproduction
in Relation to the Central Valley Project. California
Fish and Game, Vol. 34, No. 4, pp. 171-187.
1950 Striped Bass Reproduction in the Sacramento River
System in 1948. California Fish and Game, Vol. 36, No.
2, pp. 135-145.
Clark, G. H.
1929 Sacramento-San Joaquin Salmon (Oncorhynchus tshawytscha)
Fishery of California. California Fish and
Game, Fish Bulletin No. 17.
1932 The Striped Bass Supply of California, Past and Present
California Fish and Game, Vol. 18, No. 4, pp.
297-298.
1933 Fluctuations in the Abundance of Striped Bass (Roccus
lineatus) in California. California Department of Fish
and Game, Fish Bulletin No. 39.
1934 Tagging of Striped Bass. California Fish and Game,
Vol. 20, No. 1, pp. 14-19.
1936 A Second Report on Striped Bass Tagging. California
Fish and Game, Vol. 22, No. 4, pp. 272-283.
1938 Weight and Age Determination of Striped Bass. California
Fish and Game, Vol. 24, No. 2, pp. 176-177.
Cole, Charles E.
1930 Angling for Striped Bass. California Fish and Game,
Vol. 16, No. 4, pp. 286-290.
Craig, J. A.
1928 The Striped Bass Supply of California. California
Fish and Game, Vol. 14, No. 4, pp. 265-272.
1930 An Analysis of the Catch Statistics of the Striped Bass
(Roccus lineatus) Fishery of California. California
Department of Fish and Game, Fish Bulletin No. 24.
Hatton, S. Ross
1940. Progress Report on the Central Valley Fisheries Investigations,
1939. California Fish and Game, Vol.
26, No. 4, pp. 334-373.
Jackson, H. W. and R. E. Tiller
1952 Preliminary observations on spawning potential in
striped bass (Roccus saxatilis). Maryland Dept. Res.
and Ed., Pub. 93, pp. 1-6.
Johnson, W. C, and A. J. Calhoun
1952 Food Habits of California Striped Bass. California
Fish and Game, Vol. 38, No. 4, pp. 531-533.
Morgan, Alfred R. and Arthur R. Gerlach
1950 Striped Bass Studies on Coos Bay, Oregon in 1949 and
1950. Oregon Fish Commission, Contribution No. 14.
Pearson, John C.
1938 The Life History of the Striped Bass or Rockfish,
(Roccus saxatilis) (Walbaum). U. S. Department of
Commerce Bureau of Fisheries, Vol. XLIX, Bulletin
No. 28.
Raney, Edward C, Ernest F. Tresselt, Edgar H. Hollis, V. D.
Vladykov and D. H. Wallace
1952 The Striped Bass (Roccus saxatilis). Bulletin of the
Bingham Oceanographic Collection, Vol. 14, Article 1.
Scofield, N. B.
1910 Notes on striped bass in California. Biennial Report,
Calif. Board of Fish and Game Commissioners for
1909-1910, pp. 104-109.
Shapovalov, Leo
1936 Food of the Striped Bass. California Fish and Game,
Vol. 22, No. 4, pp. 261-270.
Skinner, John E.
1955a. California State-Wide Angling Estimates for 1953.
California Fish and Game, Vol. 41, No. 1, pp. 19-32.
1955b. Observations on the Shad Gill Net Fishery in 1954.
California Department of Fish and Game, Inland Fisheries
Branch, Administrative Report 55-3.
1957a. Incidental losses of Striped Bass in the Sacramento
River Gill Net Fisheries for Shad and Salmon. California
Department of Fish and Game, Inland Fisheries
Branch, Administrative Report 57-2.
1957b. Status of the Striped Bass—Sturgeon Study and Suggestions
for its Future. California Department of Fish
Fish and Game, Inland Fisheries Branch, Administrative
Report No. 57-11.
Smith, Hugh M.
1895 The Striped Bass History and Results of Introduction.
U. S. Fish Commission Bulletin, Vol. 15, pp. 449-458.
Woodhull, Chester
1947 Spawning Habits of the Striped Bass (Roccus saxatilis)
in California Waters. California Fish and Game,
Vol. 33, No. 2, pp. 97-101
The entire document is available in Adobe PDF format. John E. Skinner. 1962. An Historical Review of the Fish and Wildlife Resources of the San Francisco Bay Area. http://www.estuaryarchive.org/archive/skinner_1962/
San Francisco Estuary and Watershed Archive
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